Ralf Janssen
Researcher at Department of Earth Sciences; Palaeobiology
- E-mail:
- ralf.janssen@geo.uu.se
- Visiting address:
- Geocentrum, Villavägen 16
752 36 Uppsala - Postal address:
- Villavägen 16
752 36 UPPSALA
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Publications
Recent publications
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Part of Developmental Dynamics, p. 671-697, 2026
- DOI for Single‐cell sequencing reveals potential novel insights into appendage‐patterning and joint‐development in a spider
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Part of BMC Ecology and Evolution, 2026
- DOI for Conserved and diverged embryonic expression patterns of panarthropod NK-Cluster genes and new evidence for CRE-shuffling
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Part of Open Biology, 2026
- DOI for Pre-cheliceral region patterning in a spider provides new insights into the development and evolution of arthropod neurosecretory centres
- Download full text (pdf) of Pre-cheliceral region patterning in a spider provides new insights into the development and evolution of arthropod neurosecretory centres
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Part of Genome Biology and Evolution, 2025
- DOI for Assessing the Impact of Whole Genome Duplication on Gene Expression and Regulation During Arachnid Development
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Part of DISCOVER DEVELOPMENTAL BIOLOGY, 2025
- DOI for Early expression of chelicerate hedgehog orthologs and its bearing on the homology of arthropod head segments
- Download full text (pdf) of Early expression of chelicerate hedgehog orthologs and its bearing on the homology of arthropod head segments
All publications
Articles in journal
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Part of Developmental Dynamics, p. 671-697, 2026
- DOI for Single‐cell sequencing reveals potential novel insights into appendage‐patterning and joint‐development in a spider
- Download full text (pdf) of Single‐cell sequencing reveals potential novel insights into appendage‐patterning and joint‐development in a spider
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Part of BMC Ecology and Evolution, 2026
- DOI for Conserved and diverged embryonic expression patterns of panarthropod NK-Cluster genes and new evidence for CRE-shuffling
- Download full text (pdf) of Conserved and diverged embryonic expression patterns of panarthropod NK-Cluster genes and new evidence for CRE-shuffling
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Part of Open Biology, 2026
- DOI for Pre-cheliceral region patterning in a spider provides new insights into the development and evolution of arthropod neurosecretory centres
- Download full text (pdf) of Pre-cheliceral region patterning in a spider provides new insights into the development and evolution of arthropod neurosecretory centres
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Part of Genome Biology and Evolution, 2025
- DOI for Assessing the Impact of Whole Genome Duplication on Gene Expression and Regulation During Arachnid Development
- Download full text (pdf) of Assessing the Impact of Whole Genome Duplication on Gene Expression and Regulation During Arachnid Development
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Part of DISCOVER DEVELOPMENTAL BIOLOGY, 2025
- DOI for Early expression of chelicerate hedgehog orthologs and its bearing on the homology of arthropod head segments
- Download full text (pdf) of Early expression of chelicerate hedgehog orthologs and its bearing on the homology of arthropod head segments
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Single-cell RNA sequencing of mid-to-late stage spider embryos: new insights into spider development
Part of BMC Genomics, 2024
- DOI for Single-cell RNA sequencing of mid-to-late stage spider embryos: new insights into spider development
- Download full text (pdf) of Single-cell RNA sequencing of mid-to-late stage spider embryos: new insights into spider development
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Developmental and genomic insight into the origin of the tardigrade body plan
Part of Evolution & Development, 2024
- DOI for Developmental and genomic insight into the origin of the tardigrade body plan
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Part of EvoDevo, 2024
- DOI for Single-cell sequencing suggests a conserved function of Hedgehog-signalling in spider eye development
- Download full text (pdf) of Single-cell sequencing suggests a conserved function of Hedgehog-signalling in spider eye development
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New insights into mesoderm and endoderm development, and the nature of the onychophoran blastopore
Part of Frontiers in Zoology, 2024
- DOI for New insights into mesoderm and endoderm development, and the nature of the onychophoran blastopore
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Evolution of the Spider Homeobox Gene Repertoire by Tandem and Whole Genome Duplication
Part of Molecular biology and evolution, 2023
- DOI for Evolution of the Spider Homeobox Gene Repertoire by Tandem and Whole Genome Duplication
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Expression of posterior Hox genes and opisthosomal appendage development in a mygalomorph spider
Part of Development, Genes and Evolution, p. 107-121, 2023
- DOI for Expression of posterior Hox genes and opisthosomal appendage development in a mygalomorph spider
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Comment on "The lower Cambrian lobopodian Cardiodictyon resolves the origin of euarthropod brains"
Part of Science, 2023
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Part of Developmental Dynamics, p. 172-185, 2023
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Phylogenetic analysis of forkhead transcription factors in the Panarthropoda
Part of Development, Genes and Evolution, p. 39-48, 2022
- DOI for Phylogenetic analysis of forkhead transcription factors in the Panarthropoda
- Download full text (pdf) of Phylogenetic analysis of forkhead transcription factors in the Panarthropoda
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Embryonic expression patterns of Wnt genes in the RTA-clade spider Cupiennius salei
Part of Gene Expression Patterns, 2022
- DOI for Embryonic expression patterns of Wnt genes in the RTA-clade spider Cupiennius salei
- Download full text (pdf) of Embryonic expression patterns of Wnt genes in the RTA-clade spider Cupiennius salei
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Lack of evidence for conserved parasegmental grooves in arthropods
Part of Development, Genes and Evolution, p. 27-37, 2022
- DOI for Lack of evidence for conserved parasegmental grooves in arthropods
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A comprehensive study of arthropod and onychophoran Fox gene expression patterns
Part of PLOS ONE, 2022
- DOI for A comprehensive study of arthropod and onychophoran Fox gene expression patterns
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Molecular evidence for a single origin of ultrafiltration-based excretory organs
Part of Current Biology, p. 3629-+, 2021
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Part of G3, 2021
- DOI for Widespread retention of ohnologs in key developmental gene families following whole-genome duplication in arachnopulmonates
- Download full text (pdf) of Widespread retention of ohnologs in key developmental gene families following whole-genome duplication in arachnopulmonates
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Panarthropod tiptop/teashirt and spalt orthologs and their potential role as "trunk"-selector genes
Part of EvoDevo, 2021
- DOI for Panarthropod tiptop/teashirt and spalt orthologs and their potential role as "trunk"-selector genes
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Part of EvoDevo, 2021
- DOI for A chelicerate Wnt gene expression atlas: novel insights into the complexity of arthropod Wnt-patterning
- Download full text (pdf) of A chelicerate Wnt gene expression atlas: novel insights into the complexity of arthropod Wnt-patterning
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Part of Arthropod structure & development, 2021
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Part of Development, Genes and Evolution, p. 239-245, 2020
- DOI for Expression of the zinc finger transcription factor Sp6-9 in the velvet worm Euperipatoides kanangrensis suggests a conserved role in appendage development in Panarthropoda
- Download full text (pdf) of Expression of the zinc finger transcription factor Sp6-9 in the velvet worm Euperipatoides kanangrensis suggests a conserved role in appendage development in Panarthropoda
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Part of Development, Genes and Evolution, p. 65-73, 2020
- DOI for The forkhead box containing transcription factor FoxB is a potential component of dorsal-ventral body axis formation in the spider Parasteatoda tepidariorum
- Download full text (pdf) of The forkhead box containing transcription factor FoxB is a potential component of dorsal-ventral body axis formation in the spider Parasteatoda tepidariorum
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Part of Development, Genes and Evolution, p. 247-256, 2020
- DOI for The embryonic expression pattern of a second, hitherto unrecognized, paralog of the pair-rule gene sloppy-paired in the beetle Tribolium castaneum
- Download full text (pdf) of The embryonic expression pattern of a second, hitherto unrecognized, paralog of the pair-rule gene sloppy-paired in the beetle Tribolium castaneum
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Phylogenetic analysis and embryonic expression of panarthropod Dmrt genes
Part of Frontiers in Zoology, 2019
- DOI for Phylogenetic analysis and embryonic expression of panarthropod Dmrt genes
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FoxB, a new and highly conserved key factor in arthropod dorsal-ventral (DV) limb patterning
Part of EvoDevo, 2019
- DOI for FoxB, a new and highly conserved key factor in arthropod dorsal-ventral (DV) limb patterning
- Download full text (pdf) of FoxB, a new and highly conserved key factor in arthropod dorsal-ventral (DV) limb patterning
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Embryonic expression of priapulid Wnt genes
Part of Development, Genes and Evolution, p. 125-135, 2019
- DOI for Embryonic expression of priapulid Wnt genes
- Download full text (pdf) of Embryonic expression of priapulid Wnt genes
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The last common ancestor of Ecdysozoa had an adult terminal mouth
Part of Arthropod structure & development, p. 155-158, 2019
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Gene expression analysis of potential morphogen signalling modifying factors in Panarthropoda
Part of EvoDevo, 2018
- DOI for Gene expression analysis of potential morphogen signalling modifying factors in Panarthropoda
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Part of BMC Evolutionary Biology, 2018
- DOI for Embryonic expression patterns and phylogenetic analysis of panarthropod sox genes: insight into nervous system development, segmentation and gonadogenesis
- Download full text (pdf) of Embryonic expression patterns and phylogenetic analysis of panarthropod sox genes: insight into nervous system development, segmentation and gonadogenesis
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Part of Development, Genes and Evolution, p. 171-178, 2018
- DOI for Embryonic expression of a Long Toll (Loto) gene in the onychophorans Euperipatoides kanangrensis and Cephalofovea clandestina
- Download full text (pdf) of Embryonic expression of a Long Toll (Loto) gene in the onychophorans Euperipatoides kanangrensis and Cephalofovea clandestina
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The house spider genome reveals an ancient whole-genome duplication during arachnid evolution
Part of BMC Biology, 2017
- DOI for The house spider genome reveals an ancient whole-genome duplication during arachnid evolution
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Part of EvoDevo, 2017
- DOI for Comparative analysis of gene expression patterns in the arthropod labrum and the onychophoran frontal appendages, and its implications for the arthropod head problem
- Download full text (pdf) of Comparative analysis of gene expression patterns in the arthropod labrum and the onychophoran frontal appendages, and its implications for the arthropod head problem
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Part of Developmental Biology, p. 155-164, 2017
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Origin and Evolution of the Panarthropod Head - a Deep Time Perspective
Part of Integrative and Comparative Biology, 2017
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A molecular view of onychophoran segmentation
Part of Arthropod structure & development, p. 341-353, 2017
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Gene expression reveals evidence for EGFR-dependent proximal-distal limb patterning in a myriapod
Part of Evolution & Development, p. 124-135, 2017
- DOI for Gene expression reveals evidence for EGFR-dependent proximal-distal limb patterning in a myriapod
- Download full text (pdf) of Gene expression reveals evidence for EGFR-dependent proximal-distal limb patterning in a myriapod
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Part of Development, Genes and Evolution, p. 69-77, 2016
- DOI for Gene expression analysis reveals that Delta/Notch signalling is not involved in onychophoran segmentation
- Download full text (pdf) of Gene expression analysis reveals that Delta/Notch signalling is not involved in onychophoran segmentation
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The evolution and expression of panarthropod frizzled genes
Part of Frontiers in Ecology and Evolution, 2015
- DOI for The evolution and expression of panarthropod frizzled genes
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Fate and nature of the onychophoran mouth-anus furrow and its contribution to the blastopore
Part of Proceedings of the Royal Society of London. Biological Sciences, 2015
- DOI for Fate and nature of the onychophoran mouth-anus furrow and its contribution to the blastopore
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Aspects of dorso-ventral and proximo-distal limb patterning in onychophorans
Part of Evolution & Development, p. 21-33, 2015
- DOI for Aspects of dorso-ventral and proximo-distal limb patterning in onychophorans
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Part of International Journal of Developmental Biology, p. 343-347, 2014
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Onychophoran Hox genes and the evolution of arthropod Hox gene expression
Part of Frontiers in Zoology, p. 22, 2014
- DOI for Onychophoran Hox genes and the evolution of arthropod Hox gene expression
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Part of EvoDevo, p. 14, 2014
- DOI for Analysis of the Wnt gene repertoire in an onychophoran provides new insights into the evolution of segmentation
- Download full text (pdf) of Analysis of the Wnt gene repertoire in an onychophoran provides new insights into the evolution of segmentation
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Expression of arthropod distal limb-patterning genes in the onychophoran Euperipatoides kanangrensis
Part of Development, Genes and Evolution, p. 87-96, 2014
- DOI for Expression of arthropod distal limb-patterning genes in the onychophoran Euperipatoides kanangrensis
- Download full text (pdf) of Expression of arthropod distal limb-patterning genes in the onychophoran Euperipatoides kanangrensis
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Part of Gene Expression Patterns, p. 55-61, 2014
- DOI for Identification and embryonic expression of Wnt2, Wnt4, Wnt5 and Wnt9 in the millipede Glomeris marginata (Myriapoda: Diplopoda)
- Download full text (pdf) of Identification and embryonic expression of Wnt2, Wnt4, Wnt5 and Wnt9 in the millipede Glomeris marginata (Myriapoda: Diplopoda)
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Part of Die Naturwissenschaften, p. 33-43, 2013
- DOI for Developmental abnormalities in Glomeris marginata (Villers 1789) (Myriapoda: Diplopoda): implications for body axis determination in a myriapod
- Download full text (pdf) of Developmental abnormalities in Glomeris marginata (Villers 1789) (Myriapoda: Diplopoda): implications for body axis determination in a myriapod
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Part of Developmental Biology, p. 224-234, 2013
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A curious abnormally developed embryo of the pill millipede Glomeris marginata (Villers, 1789)
Part of ZooKeys, p. 67-75, 2013
- DOI for A curious abnormally developed embryo of the pill millipede Glomeris marginata (Villers, 1789)
- Download full text (pdf) of A curious abnormally developed embryo of the pill millipede Glomeris marginata (Villers, 1789)
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Part of Development, Genes and Evolution, p. 299-309, 2012
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Part of BMC Developmental Biology, p. 15, 2012
- DOI for Expression of pair rule gene orthologs in the blastoderm of a myriapod: evidence for pair rule-like mechanisms?
- Download full text (pdf) of Expression of pair rule gene orthologs in the blastoderm of a myriapod: evidence for pair rule-like mechanisms?
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Deuterostomic Development in the Protostome Priapulus caudatus
Part of Current Biology, p. 2161-2166, 2012
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Part of BMC Evolutionary Biology, p. 50, 2011
- DOI for Expression of collier in the premandibular segment of myriapods: support for the traditional Atelocerata concept or a case of convergence?
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Expression of myriapod pair rule gene orthologs
Part of EvoDevo, 2011
- DOI for Expression of myriapod pair rule gene orthologs
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Part of Development, Genes and Evolution, p. 351-355, 2011
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Diplosegmentation in the pill millipede Glomeris marginata is the result of dorsal fusion
Part of Evolution & Development, p. 477-487, 2011
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Gene expression suggests conserved mechanisms patterning the heads of insects and myriapods
Part of Developmental Biology, p. 64-72, 2011
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Part of BMC Evolutionary Biology, p. 374, 2010
- DOI for Conservation, loss, and redeployment of Wnt ligands in protostomes: implications for understanding the evolution of segment formation
- Download full text (pdf) of Conservation, loss, and redeployment of Wnt ligands in protostomes: implications for understanding the evolution of segment formation
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Part of EvoDevo, 2010
- DOI for Gene expression suggests conserved aspects of Hox gene regulation in arthropods and provides additional support for monophyletic Myriapoda
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Part of Development, Genes and Evolution, p. 117-122, 2010
- DOI for Head patterning and Hox gene expression in an onychophoran and its implications for the arthropod head problem
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Part of Evolution & Development, p. 363-372, 2010
- DOI for Gene expression patterns in an onychophoran reveal that regionalization predates limb segmentation in pan-arthropods
- Download full text (pdf) of Gene expression patterns in an onychophoran reveal that regionalization predates limb segmentation in pan-arthropods
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Part of Insect molecular biology (Print), p. 47-62, 2010
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Genome Sequence of the Pea Aphid Acyrthosiphon pisum
Part of PLoS biology, 2010
- DOI for Genome Sequence of the Pea Aphid Acyrthosiphon pisum
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Embryonic development and the understanding of the adult body plan in myriapods
Part of Soil Organisms, p. 337-346, 2009
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The hatching larva of the priapulid worm Halicryptus spinulosus
Part of Frontiers in Zoology, 2009
- DOI for The hatching larva of the priapulid worm Halicryptus spinulosus
- Download full text (pdf) of The hatching larva of the priapulid worm Halicryptus spinulosus
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Hatching and earliest larval stages of the priapulid worm Priapulus caudatus
Part of Invertebrate biology., p. 157-171, 2009
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Early embryonic development of the priapulid worm Priapulus caudatus
Part of Evolution & Development, p. 326-338, 2008
- DOI for Early embryonic development of the priapulid worm Priapulus caudatus
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Evidence for Wg-independent tergite boundary formation in the millipede Glomeris marginata
Part of Development, Genes and Evolution, p. 361-370, 2008
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Diverged and conserved aspects of heart formation in a spider
Part of Evolution & Development, p. 155-165, 2008
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Part of Evolution & Development, p. 143-154, 2008
- DOI for The T-box genes H15 and optomotor-blind in the spiders Cupiennius salei, Tegenaria atrica and Achaearanea tepidariorum and the dorsoventral axis of arthropod appendages
- Download full text (pdf) of The T-box genes H15 and optomotor-blind in the spiders Cupiennius salei, Tegenaria atrica and Achaearanea tepidariorum and the dorsoventral axis of arthropod appendages
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A review of the correlation of tergites, sternites, and leg pairs in diplopods
Part of Frontiers in Zoology, 2006
- DOI for A review of the correlation of tergites, sternites, and leg pairs in diplopods
- Download full text (pdf) of A review of the correlation of tergites, sternites, and leg pairs in diplopods
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The ten Hox genes of the millipede Glomeris marginata
Part of Development, Genes and Evolution, p. 451-465, 2006
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Part of Norwegian Journal of Entomology, p. 129-137, 2006
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Part of Evolution & Development, p. 51-57, 2005
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Pair rule gene orthologs in spider segmentation
Part of Evolution & Development, p. 618-628, 2005
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Part of Developmental Biology, p. 89-104, 2004
- DOI for Gene expression suggests decoupled dorsal and ventral segmentation in the millipede Glomeris marginata (Myriapoda: Diplopoda)
- Download full text (pdf) of Gene expression suggests decoupled dorsal and ventral segmentation in the millipede Glomeris marginata (Myriapoda: Diplopoda)
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Part of Developmental Biology, p. 119-140, 2003
Articles, review/survey
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Hox genes in spiders: Their significance for development and evolution
Part of Seminars in Cell and Developmental Biology, p. 24-34, 2024
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Review of extra-embryonic tissues in the closest arthropod relatives, onychophorans and tardigrades
Part of Philosophical Transactions of the Royal Society of London. Biological Sciences, 2022
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Origin and evolution of the panarthropod head: A palaeobiological and developmental perspective
Part of Arthropod structure & development, p. 354-379, 2017