Palaeobiology

Paleobiology is an interdisciplinary field of research that studies the Earth and how life on Earth has developed and adapted to changes in the environment and climate over long periods of time. We also investigate the biological and geological processes that have influenced and shaped the history of life on our planet. Paleobiologists seek answers to questions such as; How have different species evolved and disappeared over time? What factors have influenced these processes? How have changes in climate affected life on Earth and its evolutionary patterns?
Description
As far as we know, our Earth is the only planet in the universe where there is life. Humans represent only a fraction of all forms of life that have existed on our planet. At the Department of Earth Sciences, we strive to understand how life has evolved over millions of years and how it has adapted to constant changes in the environment and climate. We do this by combining traditional studies of fossils with modern biological methods. Of particular interest is the development of animals in connection with major biological events, especially what happened during the so-called Cambrian explosion – a short period more than 500 million years ago when we see a rapid increase in, among other things, shell-bearing animals. But it's not just the Cambrian explosion that attracts interest. The development of more modern animal groups in the middle of the Cretaceous period, as well as the major extinction event 65 million years ago when, among other things, dinosaurs became extinct, provides additional pieces to the puzzle of life's history and is part of our own development.
Microorganisms are found everywhere, from deep seas to high mountain peaks and even inside mountains themselves. Humans are in various ways completely dependent on these single-celled organisms, and they regulate much of our own existence, for example, by producing much of the oxygen we breathe or by breaking down and recycling organic material. At the Department of Earth Sciences, we try, among other things, to understand where these important organisms originated and how they have evolved in relation to the global climate. Our studies span several billion years of development, but particular emphasis is placed on the last 600 million years of evolution – from a time when virtually the entire Earth was covered in ice to the effects of more recent climate fluctuations.
Our Research
Early evolution of animals
Most of our understanding of Paleozoic animal life comes from preserved remains of hard parts, that is, shells and other mineralized parts of animals. By studying fossil remains from all over the world the focus is on investigating and reconstructing the relationships within and between important animal phyla, above all arthropods and brachiopods.
Shell-bearing organisms represent only a small part of the total diversity and therefore we also study non-mineralized remains in the form of organic small carbonaceous microfossils (and acritarchs). These types of microfossils preserve unique soft parts of animals, which are otherwise usually completely missing as fossils. With the help of these we can fill in some of the gaps in the early animal development.
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Genetic and molecular evolution of development
Here we focus on the investigation and comparison of embryonic gene expression patterns, i.e. the place and time when genes that steer development are active. We mainly work on the so-called panarthropods, represented by the well-known arthropods (e.g. flies, butterflies, shrimps, spiders, scorpions, centipedes and millipedes), but also the enigmatic tardigrades (water bears) and the onychophorans (velvet worms) but also priapulids, vermiform ecdysozoan critters that are known since the Cambrian era, and that may share key morphological features with the last common ancestor of Ecdysozoa as a whole.

Phytoplankton evolution
Marine phytoplankton, such as diatoms and coccolithophores, are small algae that play a crucial role in marine ecosystems and the global carbon cycle. These algae sequester large amounts of the greenhouse gas carbon dioxide, just like trees and plants on land. Under current climate change, there are many questions of both scientific and societal interest: How will marine algae adapt to a much warmer world? How fast (or slow) will these adaptations occur? Will current species go extinct?
We can learn a lot about modern algae through field-based studies and laboratory experiments, but to understand the long-term effects of climate change and the evolution of phytoplankton, we need to look back in time and study how marine plankton communities have evolved on geological timescales. Scientific deep-sea drilling provides us access to a unique archive of microfossils and paleoclimate data from earlier periods of Earth's evolution. Deep-sea sediments store information about what the climate was like in the distant past, for example when only the southern hemisphere was glaciated and sea surface temperatures exceeded 28.5°C in the North Atlantic. Our research aims at understanding the dynamic interactions between climate change and marine phytoplankton across both short and long timescales, and relating this knowledge to changes in the global carbon cycle.

Biogeochemistry
Biogeochemistry explores the spatial and temporal changes in the natural environment driven by biological, geological, chemical, and physical processes. Life on Earth depends on nutrients such as carbon, nitrogen, and trace elements, and their availability for life is determined by the natural processes that circulate them through the environment. Biogeochemistry and our research focus aims to understand these circulation patterns to comprehend how life originated, developed, and adapted to environmental changes over time. We also aim to understand the abiotic processes that circulate elements in the environment, such as weathering and depositional patterns, as well as tracers of sources and sinks of elements in nature. Biogeochemistry spans the entire globe and all timescales and is an important base for a fundamental understanding of elemental circulation on our planet.

PhD Studies
According to international evaluations, Uppsala University has some of the most comprehensive research in geosciences in Europe, and our doctoral students at Uppsala University are among the most satisfied with their doctoral education. We offer doctoral studies in eight research areas.
Publications
Part of Journal of the Geological Society, 2025
- DOI for Vast sediment dispersal in an inland sea: detrital zircon geochronology of the lower Cambrian Mickwitzia Sandstone (southern Sweden) indicates a Timanian source
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Exceptionally Preserved Setae: A Possible Morphological Synapomorphy of Cambrian Lophotrochozoans
Part of Evolution & Development, 2025
- DOI for Exceptionally Preserved Setae: A Possible Morphological Synapomorphy of Cambrian Lophotrochozoans
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Part of Chemical Engineering Journal, 2025
- DOI for Unraveling the effects of temperature on mass transfer and microbiology in thermophilic and extreme thermophilic trickle bed biomethanation reactors
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Cambrian carbonaceous protoconodonts and the early fossil record of the Chaetognatha
Part of Proceedings of the Royal Society of London. Biological Sciences, 2025
- DOI for Cambrian carbonaceous protoconodonts and the early fossil record of the Chaetognatha
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Skin, scales, and cells in a Jurassic plesiosaur
Part of Current Biology, 2025
- DOI for Skin, scales, and cells in a Jurassic plesiosaur
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Part of Geobiology, 2025
- DOI for Experimental Mineralisation of a Filamentous Hydrogenotrophic Methanogen in Carbonate, Phosphate, and Silicate
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Part of Palaeoworld, 2025
Fauna of the Sæterdal Formation (Cambrian Series 2, Stage 4) of North Greenland (Laurentia)
Part of Bulletin of the Geological Society of Denmark, p. 1-13, 2025
- DOI for Fauna of the Sæterdal Formation (Cambrian Series 2, Stage 4) of North Greenland (Laurentia)
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Part of Earth-Science Reviews, 2025
- DOI for Thickness of the stratigraphic record of Britain: How the fidelity of geological and fossil data is unrelated to rock quantity
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Part of DISCOVER DEVELOPMENTAL BIOLOGY, 2025
- DOI for Early expression of chelicerate hedgehog orthologs and its bearing on the homology of arthropod head segments
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Part of PeerJ, 2025
- DOI for A new specimen of Plesiopterys wildi reveals the diversification of cryptoclidian precursors and possible endemism within European Early Jurassic plesiosaur assemblages
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Part of Earth-Science Reviews, 2025
- DOI for The Cambrian Series 2-Miaolingian boundary interval in Australia: biostratigraphic subdivision and implications for global multi-proxy correlation
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Part of Geobiology, 2025
- DOI for Troubles With Tubules: How Do Iron-Mineral Chemical Gardens Differ From Iron-Mineralized Sheaths of Iron Oxidizing Bacteria?
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Part of Australian Journal of Earth Sciences, p. 182-212, 2025
- DOI for Integrated biostratigraphy, chemostratigraphy and geochronology of the lower Cambrian succession in the western Stansbury Basin, South Australia
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Onset of strong Iceland-Scotland overflow water 3.6 million years ago
Part of Nature Communications, 2025
- DOI for Onset of strong Iceland-Scotland overflow water 3.6 million years ago
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Part of npj Materials Degradation, 2025
- DOI for Microbial dissolution of Gran Canaria lapilli in small-scale flow through columns: carbon dioxide removal potential
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Middle Cambrian (Wuliuan Stage) Small Shelly Fossils from North Greenland (Laurentia)
Part of Bulletin of Geosciences, p. 1-56, 2025
Extending the diversity of grasping spines in middle Cambrian stem-group Chaetognathifera
Part of Alcheringa, p. 17-29, 2025
- DOI for Extending the diversity of grasping spines in middle Cambrian stem-group Chaetognathifera
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Part of Chemie der Erde, 2025
- DOI for Mixed fluid processes in Fe-Mn dendrite formation and associated carbon and nickel isotope fractionation
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Part of Paleoceanography and Paleoclimatology, 2025
- DOI for Major Shifts in Equatorial Atlantic and Pacific Calcareous Nannofossil Assemblages across the Early Eocene Climatic Optimum (EECO; ~53–49 Ma)
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Palaeostomate Bryozoans from Glacial Erratics in the Tvären Region, Sweden
Part of Minerals, p. 136, 2025
- DOI for Palaeostomate Bryozoans from Glacial Erratics in the Tvären Region, Sweden
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Geomagnetic Excursions Recorded in North Atlantic IODP Expedition 395C Sites U1555 and U1563
Part of Geochemistry Geophysics Geosystems, 2025
- DOI for Geomagnetic Excursions Recorded in North Atlantic IODP Expedition 395C Sites U1555 and U1563
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Part of Geophysical Journal International, 2025
- DOI for Magnetosomal Greigite as the source of intermediate and inverse magnetic fabrics at IODP site M0061 (Baltic Sea)
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Part of Applied Soil Ecology, 2025
- DOI for Nitrogen rate drives the effectiveness of 3,4-Dimethylpyrazole phosphate (DMPP) in reducing N2O emissions from limed soil subjected to temporary waterlogging
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How earthworms thrive and drive silicate rock weathering in an artificial organo-mineral system
Part of Applied Geochemistry, 2025
Palaeontology: Straightening out early echinoderms
Part of Current Biology, 2025
Hedgehog Signaling during Gut Formation in the Freshwater Leech, Helobdella austinensis
Part of Cells Tissues Organs, p. 194-205, 2025
- DOI for Hedgehog Signaling during Gut Formation in the Freshwater Leech, Helobdella austinensis
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Part of Papers in Palaeontology, 2025
- DOI for Biostratigraphy and correlation of the Cambrian Series 2 (Stage 4) to Miaolingian (Wuliuan) Thorntonia Limestone, south-eastern Georgina Basin (Queensland, Australia)
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Part of Biology, 2025
- DOI for Branched Setae or Attached Macroalgae: A Case Study of an Exceptionally Preserved Brachiopod from the Early Cambrian Chengjiang Lagerstätte
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Part of Lethaia, p. 1-15, 2025
- DOI for First report of epithelial cell moulds with nuclei impressions preserved in lower Cambrian (Stage 3) brachiopods from South China
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New insights into mesoderm and endoderm development, and the nature of the onychophoran blastopore
Part of Frontiers in Zoology, 2024
- DOI for New insights into mesoderm and endoderm development, and the nature of the onychophoran blastopore
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2024
Part of eLIFE, 2024
- DOI for Evolution and diversity of biomineralized columnar architecture in early Cambrian phosphatic-shelled brachiopods
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Tergomyan molluscs from the Early Ordovician of the Llangynog Inlier, South Wales, UK
Part of Palaeontologische Zeitschrift, p. 17-28, 2024
- DOI for Tergomyan molluscs from the Early Ordovician of the Llangynog Inlier, South Wales, UK
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Part of Bulletin of the Geological Society of Denmark, p. 57-66, 2024
- DOI for Euendolith borings in Chancelloria and Nisusia from the middle Cambrian (Miaolingian) of North Greenland (Laurentia)
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Part of NPJ MATERIALS DEGRADATION, 2024
- DOI for Organic carbon source controlled microbial olivine dissolution in small-scale flow-through bioreactors, for CO2 removal
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Part of Aquaculture, 2024
- DOI for Yarrowia lipolytica yeast as a dietary supplement for rainbow trout (Oncorhynchus mykiss): Effects on gut microbiota, health and immunity
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Part of Frontiers in Microbiology, 2024
- DOI for Catabolism and interactions of syntrophic propionate- and acetate oxidizing microorganisms under mesophilic, high-ammonia conditions
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Impact of additives on syntrophic propionate and acetate enrichments under high-ammonia conditions
Part of Applied Microbiology and Biotechnology, 2024
- DOI for Impact of additives on syntrophic propionate and acetate enrichments under high-ammonia conditions
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Controls on Li partitioning and isotopic fractionation in inorganic calcite
Part of Geochimica et Cosmochimica Acta, p. 91-102, 2024
- DOI for Controls on Li partitioning and isotopic fractionation in inorganic calcite
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Part of EvoDevo, 2024
- DOI for Single-cell sequencing suggests a conserved function of Hedgehog-signalling in spider eye development
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Part of Systematic Biology, p. 223-234, 2024
- DOI for Two Notorious Nodes: A Critical Examination of Relaxed Molecular Clock Age Estimates of the Bilaterian Animals and Placental Mammals
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Helcionelloid molluscs from the lower Cambrian (Series 2, Stage 4) of southern Sweden
Part of Historical Biology, p. 1854-1882, 2024
- DOI for Helcionelloid molluscs from the lower Cambrian (Series 2, Stage 4) of southern Sweden
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Developmental and genomic insight into the origin of the tardigrade body plan
Part of Evolution & Development, 2024
- DOI for Developmental and genomic insight into the origin of the tardigrade body plan
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Part of ICES Journal of Marine Science, p. 1137-1149, 2024
- DOI for Age validation of yellowfin and bigeye tuna using post-peak bomb radiocarbon dating confirms long lifespans in the western and central Pacific Ocean
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Lithostratigraphy of the Portfjeld Group (Ediacaran - lowermost Cambrian) of North Greenland
Part of GEUS Bulletin, 2024
- DOI for Lithostratigraphy of the Portfjeld Group (Ediacaran - lowermost Cambrian) of North Greenland
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Multiple cis-regulatory elements control prox1a expression in distinct lymphatic vascular beds
Part of Development, 2024
Problematic tubular fossils from the Portfjeld Formation (Ediacaran) of North Greenland
Part of Journal of Paleontology, p. 221-231, 2024
- DOI for Problematic tubular fossils from the Portfjeld Formation (Ediacaran) of North Greenland
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Unusual diagenesis of Cambrian chancelloriids from Greenland and Siberia
Part of Alcheringa, p. 419-429, 2024
- DOI for Unusual diagenesis of Cambrian chancelloriids from Greenland and Siberia
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Periostracum in Cambrian helcionelloid and rostroconch molluscs: comparison to modern taxa
Part of Lethaia, p. 1-17, 2024
- DOI for Periostracum in Cambrian helcionelloid and rostroconch molluscs: comparison to modern taxa
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