The two-year programme is not given in its entirety in Uppsala but jointly with three other universities. In each cohort, about half of the students start in Uppsala and the rest in Groningen. During the second semester, the studies are conducted either in Montpellier or in Munich. During semester 3 and 4, students do their own research projects at one or some of the four partner universities or at the affiliated universities.

Level/Depth is given with the abbreviations:

A1N - Second cycle, has only first-cycle courses as entry requirements, at least 120 credits

A1F - Second cycle, has second-cycle courses as entry requirements

A2E - Second cycle, degree project for Master of Arts/Master of Science (120 credits)

Main field of study is indicated according to the following abbreviations:

B = Biology

Level/Depth and Main field of study refers to the classification done by Uppsala University, also for courses provided by the other universities.

Semester 1

Track - Uppsala University

Period 1

• Evolutionary Processes, 15 credits (1BG373) Main field(s) of study and in-depth level: Biology A1N

Period 2

• Evolutionary Patterns, 15 credits (1BG306) Main field(s) of study and in-depth level: Biology A1F

Track - RuG - Rijksuniversität Groningen

Period	Course code	Course name	Credits	Level/Depth	Main field of study
1	WMEV003-10	Behaviour, Ecology & Evolution	10	A1N	B
1	WMEV006-05	Evolutionary Theory	5	A1N	B
2	WMMB005-05	Principles of Population Genetics in Natural Populations	5	A1N	B
2	WMEV011-05	Genomics in Ecology and Evolution	5	A1N	B
2	WMEV012-05	Research Proposal Ecology and Evolution	5	A1N	B

Semester 2

Spår - UM - University of Montpellier

Period	Course code	Course name	Credits	Level/Depth	Main field of study
3	HAB816B	Population genetics	3	A1N	B
3	HAB820B	Statistics	3	A1N	B
3	HAB809B	Genetic Data analysis	3	A1N	B
3	HAB811B	Hot topics in Ecology & Evolution: presenting and discussing selected research	2	A1N	B
3	HAB813B	Modelling	3	A1N	B
4	HAB807B	Evolutionary Applications	1	A1N	B
4	HAB819B	Research project	15	A1N	B

Track - LMU - Ludwig Maximilians Universität München

Period	Course code	Course name	Credits	Level/Depth	Main field of study
3	LSF19465	Basic Evolutionary Genomics	3	A1N	B
4	LSF19463	Advanced Evolutionary Genomics	3	A1N	B
4	LSF19435	Statistics	3	A1N	B
LMU	LSF19451	EES Excursion	3	A1N	B

Comment: The second semester in Munich begins around mid-April. While at LMU, MEME students will mostly participate in the courses offered in the master's program in Evolution, Ecology and Systematics (EES). However, they can also participate in courses that are offered in the other master´s programs of the LMU Faculty of Biology in Plant Sciences (PLS), in Molecular and Cellular Biology (MCB) or Human Biology (MHB).  All courses are taught in English.

Semester 3

During semester 3 and 4 the students choose at which university they want to study. It is also possible to do a project at Harvard University or at Université Lausanne.

Period	Course code	Course name	Credits	Level/Depth	Main field of study
1-2	RuG	Thesis project	30	A2E	B
1-2	UM2	Thesis project	30	A2E	B
1-2	LMU	Thesis project	30	A2E	B

• Degree Project in Biology, 30 credits (1BG362) Main field(s) of study and in-depth level: Biology A2E
• Project Work in Biology, 30 credits (1BG368) Main field(s) of study and in-depth level: Biology A1N

Semester 4

During semester 3 and 4 the students choose at which university they want to study. It is also possible to do a project at Harvard University or at Université Lausanne.

Period	Course code	Course name	Credits	Level/Depth	Main field of study
3-4	RuG	Thesis project	30	A2E	B
3-4	UM2	Thesis project	30	A2E	B
3-4	LMU	Project work	30	A1F	B

• Project Work in Biology, 30 credits (1BG368) Main field(s) of study and in-depth level: Biology A1N
• Degree Project in Biology, 30 credits (1BG362) Main field(s) of study and in-depth level: Biology A2E