Lars Holmer
Professor in Historical geology and Palaeontology at Department of Earth Sciences; Palaeobiology
- Telephone:
- +46 18 471 27 61
- Mobile phone:
- +46 70 468 26 55
- E-mail:
- Lars.Holmer@pal.uu.se
- Visiting address:
- Geocentrum, Villavägen 16
752 36 Uppsala - Postal address:
- Villavägen 16
752 36 UPPSALA
Download contact information for Lars Holmer at Department of Earth Sciences; Palaeobiology
Director at Campus Management Organisation; Campus Lagerträdet
- Mobile phone:
- +46 70 468 26 55
- E-mail:
- lars.holmer@pal.uu.se
- Visiting address:
- Reception
Norbyvägen 14 - Postal address:
- Norbyvägen 18D
752 36 UPPSALA
Download contact information for Lars Holmer at Campus Management Organisation; Campus Lagerträdet
- Academic merits:
- Docent
- CV:
- Download CV
- ORCID:
- 0000-0003-3629-0049
Short presentation
The main aim of my research is to resolve the origin and earliest phylogeny of the brachiopods and other lophophorates, by detailed palaeobiological investigation of their oldest remains in the fossil record in combination with knowledge from molecular and morphological/anatomical studies of the living forms.
Media
Podcast on brachiopods
https://www.palaeocast.com/episode-40-brachiopods/
Lecture for Geological Association
Mickwitzia monilifera and Cambrian Brachiopod Phylogeny
https://media.medfarm.uu.se/play/video/9378
Brachiopods' life strategies in several key geological periods.
Movie illustrating the article https://www.cell.com/current-biology/fulltext/S0960-9822(23)00178-1

Publications
Recent publications
Part of Communications Biology, 2025
- DOI for Combining soft-bodied and three-dimensional fossils to reveal evolutionary modifications in early lingulellotretid brachiopods
- Download full text (pdf) of Combining soft-bodied and three-dimensional fossils to reveal evolutionary modifications in early lingulellotretid brachiopods
Part of Lethaia, p. 1-15, 2025
- DOI for First report of epithelial cell moulds with nuclei impressions preserved in lower Cambrian (Stage 3) brachiopods from South China
- Download full text (pdf) of First report of epithelial cell moulds with nuclei impressions preserved in lower Cambrian (Stage 3) brachiopods from South China
Part of Biology, 2025
- DOI for Branched Setae or Attached Macroalgae: A Case Study of an Exceptionally Preserved Brachiopod from the Early Cambrian Chengjiang Lagerstätte
- Download full text (pdf) of Branched Setae or Attached Macroalgae: A Case Study of an Exceptionally Preserved Brachiopod from the Early Cambrian Chengjiang Lagerstätte
Palaeostomate Bryozoans from Glacial Erratics in the Tvären Region, Sweden
Part of Minerals, p. 136, 2025
- DOI for Palaeostomate Bryozoans from Glacial Erratics in the Tvären Region, Sweden
- Download full text (pdf) of Palaeostomate Bryozoans from Glacial Erratics in the Tvären Region, Sweden
Part of Palaeoworld, 2025
All publications
Articles in journal
Part of Communications Biology, 2025
- DOI for Combining soft-bodied and three-dimensional fossils to reveal evolutionary modifications in early lingulellotretid brachiopods
- Download full text (pdf) of Combining soft-bodied and three-dimensional fossils to reveal evolutionary modifications in early lingulellotretid brachiopods
Part of Lethaia, p. 1-15, 2025
- DOI for First report of epithelial cell moulds with nuclei impressions preserved in lower Cambrian (Stage 3) brachiopods from South China
- Download full text (pdf) of First report of epithelial cell moulds with nuclei impressions preserved in lower Cambrian (Stage 3) brachiopods from South China
Part of Biology, 2025
- DOI for Branched Setae or Attached Macroalgae: A Case Study of an Exceptionally Preserved Brachiopod from the Early Cambrian Chengjiang Lagerstätte
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Palaeostomate Bryozoans from Glacial Erratics in the Tvären Region, Sweden
Part of Minerals, p. 136, 2025
- DOI for Palaeostomate Bryozoans from Glacial Erratics in the Tvären Region, Sweden
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Part of Palaeoworld, 2025
Exceptionally Preserved Setae: A Possible Morphological Synapomorphy of Cambrian Lophotrochozoans
Part of Evolution & Development, 2025
- DOI for Exceptionally Preserved Setae: A Possible Morphological Synapomorphy of Cambrian Lophotrochozoans
- Download full text (pdf) of Exceptionally Preserved Setae: A Possible Morphological Synapomorphy of Cambrian Lophotrochozoans
Part of Journal of Asian Earth Sciences, 2024
- DOI for Late Ordovician lingulid brachiopods from the Pingliang Formation (Shaanxi Province, North China): Morphological and ecological implications
- Download full text (pdf) of Late Ordovician lingulid brachiopods from the Pingliang Formation (Shaanxi Province, North China): Morphological and ecological implications
Evolution of brachiopod symbiosis in the early Paleozoic
Part of Historical Biology, p. 1274-1294, 2024
Part of Gondwana Research, p. 126-141, 2024
- DOI for First multi-proxy chronostratigraphy of the lower Cambrian Byrd Group, Transantarctic Mountains and correlation within East Gondwana
- Download full text (pdf) of First multi-proxy chronostratigraphy of the lower Cambrian Byrd Group, Transantarctic Mountains and correlation within East Gondwana
Part of eLIFE, 2024
- DOI for Evolution and diversity of biomineralized columnar architecture in early Cambrian phosphatic-shelled brachiopods
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Part of Papers in Palaeontology, 2023
Evolutionary contingency in lingulid brachiopods across mass extinctions
Part of Current Biology, p. 1565-1572000, 2023
- DOI for Evolutionary contingency in lingulid brachiopods across mass extinctions
- Download full text (pdf) of Evolutionary contingency in lingulid brachiopods across mass extinctions
Part of Palaios, p. 240-245, 2023
Part of Estonian journal of earth sciences, p. 140-140, 2023
- DOI for When lingulid brachiopods became infaunal(?): Perspectives from the morphological and anatomical information
- Download full text (pdf) of When lingulid brachiopods became infaunal(?): Perspectives from the morphological and anatomical information
Part of Biology, 2023
- DOI for First Report of Small Skeletal Fossils from the Upper Guojiaba Formation (Series 2, Cambrian), Southern Shaanxi, South China
- Download full text (pdf) of First Report of Small Skeletal Fossils from the Upper Guojiaba Formation (Series 2, Cambrian), Southern Shaanxi, South China
Part of Estonian journal of earth sciences, 2023
- DOI for Reconstruction of the first consumer-driven marine ecosystem on Earth, perspectives from early Cambrian small skeletal fossils from China
- Download full text (pdf) of Reconstruction of the first consumer-driven marine ecosystem on Earth, perspectives from early Cambrian small skeletal fossils from China
Redescription and systematic position of Diandongia pista from the Chengjiang Lagerstätte
Part of Palaeoworld, p. 373-384, 2023
Cambrian (Stage 4 to Wuliuan) brachiopods from Sonora, Mexico
Part of Journal of Paleontology, p. 1-21, 2022
- DOI for Cambrian (Stage 4 to Wuliuan) brachiopods from Sonora, Mexico
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Part of Scientific Reports, 2022
- DOI for Submarine metalliferous carbonate mounds in the Cambrian of the Baltoscandian Basin induced by vent networks and water column stratification
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Part of Journal of Paleontology, p. 61-80, 2022
- DOI for Brachiopods from the Latham Shale Lagerstätte (Cambrian Series 2, Stage 4) and Cadiz Formation (Miaolingian, Wuliuan), California
- Download full text (pdf) of Brachiopods from the Latham Shale Lagerstätte (Cambrian Series 2, Stage 4) and Cadiz Formation (Miaolingian, Wuliuan), California
Part of Precambrian Research, 2022
- DOI for Late Ediacaran cavity-dwelling filamentous microorganisms accommodated in a valve-like organism from the uppermost Dengying Formation in eastern Yunnan of South China
- Download full text (pdf) of Late Ediacaran cavity-dwelling filamentous microorganisms accommodated in a valve-like organism from the uppermost Dengying Formation in eastern Yunnan of South China
Diverse endobiotic symbiont fauna from the late Katian (Late Ordovician) of Estonia
Part of Palaeontologia Electronica, 2022
- DOI for Diverse endobiotic symbiont fauna from the late Katian (Late Ordovician) of Estonia
- Download full text (pdf) of Diverse endobiotic symbiont fauna from the late Katian (Late Ordovician) of Estonia
Fossil brachiopod identification using a new deep convolutional neural network
Part of Gondwana Research, p. 290-298, 2022
Early-middle Cambrian stratigraphy and faunas from northern Siberia
Part of Acta Palaeontologica Polonica, p. 341-464, 2022
- DOI for Early-middle Cambrian stratigraphy and faunas from northern Siberia
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Part of Acta Palaeontologica Polonica, p. 207-229, 2021
- DOI for Camenellan tommotiids from the Cambrian Series 2 of East Antarctica: Biostratigraphy, palaeobiogeography, and systematics
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The oldest Cambrian trilobite - brachiopod association in South China
Part of Gondwana Research, p. 147-167, 2021
Part of Minerals, 2021
- DOI for First Report of Small Shelly Fossils from the Cambrian Miaolingian Limestones (Zhangxia and Hsuzhuang Formations) in Yiyang County, Henan Province of North China
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Part of Journal of Paleontology, p. 497-526, 2021
- DOI for Early Cambrian (Stage 4) brachiopods from the Shipai Formation in the Three Gorges area of South China
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Part of Journal of Systematic Palaeontology, p. 191-219, 2021
Part of Journal of Analytical and Applied Pyrolysis, 2021
Part of Palaeogeography, Palaeoclimatology, Palaeoecology, 2021
- DOI for Burrows filled with faecal pellets from the Cambrian (Stage 4) Guanshan biota of South China and their palaeoecological implications
- Download full text (pdf) of Burrows filled with faecal pellets from the Cambrian (Stage 4) Guanshan biota of South China and their palaeoecological implications
Part of Journal of Materials Science, p. 19884-19898, 2021
- DOI for Biomacromolecules in recent phosphate-shelled brachiopods: identification and characterization of chitin matrix
- Download full text (pdf) of Biomacromolecules in recent phosphate-shelled brachiopods: identification and characterization of chitin matrix
Part of Journal of Paleontology, p. 40-55, 2021
Part of Historical Biology, p. 3579-3584, 2021
- DOI for Possible drill holes and pseudoborings in obolid shells from the Cambrian/Ordovician boundary beds of Estonia and the uppermost Cambrian of NW Russia
- Download full text (pdf) of Possible drill holes and pseudoborings in obolid shells from the Cambrian/Ordovician boundary beds of Estonia and the uppermost Cambrian of NW Russia
Go large or go conical: allometric trajectory of an early Cambrian acrotretide brachiopod
Part of Palaeontology, p. 727-741, 2021
- DOI for Go large or go conical: allometric trajectory of an early Cambrian acrotretide brachiopod
- Download full text (pdf) of Go large or go conical: allometric trajectory of an early Cambrian acrotretide brachiopod
Part of Journal of Systematic Palaeontology, p. 1569-1588, 2020
- DOI for Ontogeny and evolutionary significance of a new acrotretide brachiopod genus from Cambrian Series 2 of South China
- Download full text (pdf) of Ontogeny and evolutionary significance of a new acrotretide brachiopod genus from Cambrian Series 2 of South China
Linguliform brachiopods from the Cambrian (Guzhangian) Karpinsk Formation of Novaya Zemlya
Part of Papers in Palaeontology, p. 571-592, 2020
- DOI for Linguliform brachiopods from the Cambrian (Guzhangian) Karpinsk Formation of Novaya Zemlya
- Download full text (pdf) of Linguliform brachiopods from the Cambrian (Guzhangian) Karpinsk Formation of Novaya Zemlya
Part of Journal of Systematic Palaeontology, p. 945-973, 2020
Part of Lethaia, p. 574-587, 2020
- DOI for Shell structure, ornamentation and affinity of the problematic early Cambrian brachiopod Heliomedusa orienta
- Download full text (pdf) of Shell structure, ornamentation and affinity of the problematic early Cambrian brachiopod Heliomedusa orienta
Part of Estonian journal of earth sciences, p. 143-153, 2020
- DOI for The Early Devonian (Emsian) acrotretid microbrachiopod Opsiconidion minor Popov, 1981, from the Alaska/Yukon Territory border and Novaya Zemlya
- Download full text (pdf) of The Early Devonian (Emsian) acrotretid microbrachiopod Opsiconidion minor Popov, 1981, from the Alaska/Yukon Territory border and Novaya Zemlya
Part of Papers in Palaeontology, p. 349-383, 2020
Part of Palaeoworld, p. 512-533, 2020
Part of RSC Advances, p. 38456-38467, 2020
- DOI for Characterization of organophosphatic brachiopod shells: spectroscopic assessment of collagen matrix and biomineral components
- Download full text (pdf) of Characterization of organophosphatic brachiopod shells: spectroscopic assessment of collagen matrix and biomineral components
Part of Science Bulletin, p. 1543-1546, 2020
- DOI for First report of brachiopods with soft parts from the Lower Cambrian Latham Shale (Series 2, Stage 4), California
- Download full text (pdf) of First report of brachiopods with soft parts from the Lower Cambrian Latham Shale (Series 2, Stage 4), California
Part of Estonian journal of earth sciences, p. 206-224, 2019
- DOI for The problematic lingulate brachiopod Aulonotreta from the Ordovician (Dapingian-Darriwilian) of Baltoscandia
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Cambrian rhynchonelliform nisusioid brachiopods: phylogeny and distribution
Part of Papers in Palaeontology, p. 559-575, 2019
- DOI for Cambrian rhynchonelliform nisusioid brachiopods: phylogeny and distribution
- Download full text (pdf) of Cambrian rhynchonelliform nisusioid brachiopods: phylogeny and distribution
Part of Scientific Reports, 2019
- DOI for Glendonite occurrences in the Tremadocian of Baltica: first Early Palaeozoic evidence of massive ikaite precipitation at temperate latitude
- Download full text (pdf) of Glendonite occurrences in the Tremadocian of Baltica: first Early Palaeozoic evidence of massive ikaite precipitation at temperate latitude
Part of Journal of Paleontology, p. 437-459, 2019
- DOI for Mollusks from the upper Shackleton Limestone (Cambrian Series 2), Central Transantarctic Mountains, East Antarctica
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Part of Lethaia, p. 424-432, 2018
Part of Journal of Paleontology, p. 33-39, 2018
Part of BMC Evolutionary Biology, 2018
- DOI for Earliest ontogeny of early Cambrian acrotretoid brachiopods: first evidence for metamorphosis and its implications
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Part of Palaeontology, p. 183-207, 2018
- DOI for Post-metamorphic allometry in the earliest acrotretoid brachiopods from the lower Cambrian (Series 2) of South China, and its implications
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Brachiopods: origin and early history
Part of Palaeontology, p. 609-631, 2017
Part of Papers in Palaeontology, p. 387-399, 2017
Part of Papers in Palaeontology, p. 317-361, 2017
- DOI for Ecology, biofacies, biogeography and systematics of micromorphic lingulate brachiopods from the Ordovician (Darriwilian–Sandbian) of south-central China
- Download full text (pdf) of Ecology, biofacies, biogeography and systematics of micromorphic lingulate brachiopods from the Ordovician (Darriwilian–Sandbian) of south-central China
Gene Expression Patterns in Brachiopod Larvae Refute the "Brachiopod-Fold" Hypothesis
Part of Frontiers in Cell and Developmental Biology, p. 1-3, 2017
- DOI for Gene Expression Patterns in Brachiopod Larvae Refute the "Brachiopod-Fold" Hypothesis
- Download full text (pdf) of Gene Expression Patterns in Brachiopod Larvae Refute the "Brachiopod-Fold" Hypothesis
Part of Acta Palaeontologica Sinica, p. 483-503, 2017
Part of Journal of Paleontology, p. 911-918, 2017
Do brachiopods show substrate-related phenotypic variation?: A case study from the Burgess Shale
Part of Palaeontology, p. 269-279, 2017
Part of Palaeoworld, p. 333-355, 2016
Part of Australasian Palaeontological Memoirs, p. 31-50, 2016
Part of GFF, p. 519-525, 2016
Part of Stratigraphy, p. 55-61, 2015
The early Cambrian tommotiid Kulparina rostrata from South Australia
Part of Journal of Paleontology, p. 920-932, 2015
Part of Stratigraphy, p. 22-30, 2015
Part of BMC Evolutionary Biology, 2015
- DOI for Competition and mimicry: the curious case of chaetae in brachiopods from the middle Cambrian Burgess Shale
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Biostratigraphy of the Cambrian–Ordovician boundary beds at Kopet-Dagh, Iran
Part of Stratigraphy, p. 40-47, 2015
Part of Annales de Paléontologie, p. 167-177, 2015
- DOI for First report of linguloid brachiopods with soft parts from the lower Cambrian (Series 2, Stage 4) of the Three Gorges area, South China
- Download full text (pdf) of First report of linguloid brachiopods with soft parts from the lower Cambrian (Series 2, Stage 4) of the Three Gorges area, South China
The new stem-group brachiopod Oymurania from the lower Cambrian of Siberia
Part of Acta Palaeontologica Polonica, p. 963-980, 2015
- DOI for The new stem-group brachiopod Oymurania from the lower Cambrian of Siberia
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Part of GFF, p. 1-8, 2015
- DOI for Taxonomy, morphology, shell structure and early ontogeny of Pelmanotreta nom. nov. from the lower Cambrian of Siberia
- Download full text (pdf) of Taxonomy, morphology, shell structure and early ontogeny of Pelmanotreta nom. nov. from the lower Cambrian of Siberia
Exceptionally preserved Mickwitzia from the Indian Springs Lagerstätte (Cambrian Stage 3), Nevada
Part of Journal of Paleontology, p. 933-955, 2015
Oldest mickwitziid brachiopod from the Terreneuvian of southern France
Part of Acta Palaeontologica Polonica, p. 755-768, 2015
- DOI for Oldest mickwitziid brachiopod from the Terreneuvian of southern France
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Himalayan Cambrian brachiopods
Part of Papers in Palaeontology, p. 345-399, 2015
Part of Palaeontology, p. 417-446, 2014
- DOI for Paterimitra pyramidalis from South Australia: Scleritome, shell structure and evolution of a lower Cambrian stem group brachiopod
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An early Cambrian agglutinated tubular lophophorate with brachiopod characters
Part of Scientific Reports, p. 4682, 2014
- DOI for An early Cambrian agglutinated tubular lophophorate with brachiopod characters
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Brachiopods hitching a ride: an early case of commensalism in the middle Cambrian Burgess Shale
Part of Scientific Reports, p. 6704, 2014
- DOI for Brachiopods hitching a ride: an early case of commensalism in the middle Cambrian Burgess Shale
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Part of Palaeogeography, Palaeoclimatology, Palaeoecology, p. 373-384, 2014
- DOI for Adaptive strategies and environmental significance of lingulid brachiopods across the late Permian extinction
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Part of GFF, p. 30-33, 2014
- DOI for New U-Pb zircon ages of the Sandbian (Upper Ordovician) "Big K-bentonite" in Baltoscandia (Estonia and Sweden) by LA-ICPMS
- Download full text (pdf) of New U-Pb zircon ages of the Sandbian (Upper Ordovician) "Big K-bentonite" in Baltoscandia (Estonia and Sweden) by LA-ICPMS
Part of Acta Palaeontologica Polonica, p. 359-365, 2014
- DOI for The problematic early Cambrian fossil Tumulduria incomperta represents the detached ventral interarea of a paterinid brachiopod
- Download full text (pdf) of The problematic early Cambrian fossil Tumulduria incomperta represents the detached ventral interarea of a paterinid brachiopod
Ordovician–Silurian Chileida—first post-Cambrian records of an enigmatic group of Brachiopoda
Part of Journal of Paleontology, p. 488-496, 2014
- DOI for Ordovician–Silurian Chileida—first post-Cambrian records of an enigmatic group of Brachiopoda
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Part of GFF, p. 539-547, 2014
- DOI for Oldest glosselline linguliform brachiopod with soft parts from the Lower Cambrian of Yunnan, Southern China
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Silurian craniide brachiopods from Gotland
Part of Palaeontology, p. 1029-1044, 2013
- DOI for Silurian craniide brachiopods from Gotland
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The oldest brachiopods from the lower Cambrian of South Australia
Part of Acta Palaeontologica Polonica, p. 93-109, 2013
- DOI for The oldest brachiopods from the lower Cambrian of South Australia
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Part of Zoomorphology, p. 379-387, 2013
- DOI for Metamorphosis in Craniiformea revisited: Novocrania anomala shows delayed development of the ventral valve
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A sclerite-bearing stem group entoproct from the early Cambrian and its implications
Part of Scientific Reports, p. 1066, 2013
- DOI for A sclerite-bearing stem group entoproct from the early Cambrian and its implications
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Cambrian (Furongian) rhynchonelliform brachiopods from the Eastern Alborz Mountains, Iran
Part of Bulletin of Geosciences, p. 525-538, 2013
- DOI for Cambrian (Furongian) rhynchonelliform brachiopods from the Eastern Alborz Mountains, Iran
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Part of Palaeontology, p. 961-970, 2013
- DOI for Morphology, ontogeny and affinities of the Hirnantian triplisiid brachiopod Streptis undifera from Baltoscandia
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Part of Science Foundation in China, p. 66-80, 2013
Part of Norwegian Journal of Geology, p. 167-174, 2012
Part of Palaeogeography, Palaeoclimatology, Palaeoecology, p. 60-67, 2012
Earliest ontogeny of Early Palaeozoic Craniiformea: compelling evidence for lecithotrophy
Part of Lethaia, p. 566-573, 2012
An early Cambrian chileate brachiopod from South Australia and its pylogenetic significance
Part of Memoirs of the Association of Australasian Palaeontologists, p. 289-294, 2011
Part of Palaeontology, p. 235-239, 2011
Part of Palaeontology, p. 253-286, 2011
Part of Palaeogeography, Palaeoclimatology, Palaeoecology, p. 206-212, 2011
Part of Geotechnique, p. 967-970, 2011
Taxonomy and biostratigraphy of Ordovician brachiopods from northeastern Ny Friesland, Spitsbergen
Part of Zootaxa, p. 1-122, 2011
Part of Journal of Paleontology, p. 460-463, 2011
Lower palaeozoic stratigraphy of Murchisonfjorden and Sparreneset, Nordaustlandet, Svalbard
Part of Geografiska Annaler. Series A, Physical Geography, p. 209-226, 2011
Part of Lethaia, p. 490-495, 2011
Part of GFF, p. 117-122, 2010
- DOI for Setatella significans, a new name for mickwitziid stem group brachiopods from the lower Cambrian of Greenland and Labrador
- Download full text (pdf) of Setatella significans, a new name for mickwitziid stem group brachiopods from the lower Cambrian of Greenland and Labrador
Earliest ontogeny of Early Palaeozoic Craniiformea: implications for brachiopod phylogeny
Part of Lethaia, p. 323-333, 2010
- DOI for Earliest ontogeny of Early Palaeozoic Craniiformea: implications for brachiopod phylogeny
- Download full text (pdf) of Earliest ontogeny of Early Palaeozoic Craniiformea: implications for brachiopod phylogeny
Part of Acta Palaeontologica Polonica, p. 495-505, 2010
- DOI for Soft-part preservation in a linguliform brachiopod from the lower Cambrian Wulongqing Formation (Guanshan Fauna) of Yunnan, South China
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Diversity fluctuations and biogeography of Ordovician brachiopod faunas in northeastern Spitsbergen
Part of Bulletin of Geosciences, p. 497-504, 2010
- DOI for Diversity fluctuations and biogeography of Ordovician brachiopod faunas in northeastern Spitsbergen
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Part of Acta Zoologica, p. 150-162, 2010
- DOI for First record of the brachiopod Lingulella waptaensis with pedicle from the Middle Cambrian Burgess Shale
- Download full text (pdf) of First record of the brachiopod Lingulella waptaensis with pedicle from the Middle Cambrian Burgess Shale
First report of the early Cambrian stem group brachiopod Mickwitzia from East Gondwana
Part of Gondwana Research, p. 145-150, 2009
Part of Palaeontologische Zeitschrift, p. 255-266, 2009
The enigmatic early cambrian salanygolina: a stem group of rhynchonelliform chileate brachiopods?
Part of Palaeontology, p. 1-10, 2009
Part of Geobios, p. 649-661, 2009
Homologous skeletal secretion in tommotiids and brachiopods
Part of Geology, p. 1143-1146, 2009
Part of Gondwana Research, p. 151-161, 2009
The scleritome of Paterimitra: an Early Cambrian stem group brachiopod from South Australia
Part of Proceedings of the Royal Society of London. Biological Sciences, p. 1651-1656, 2009
Furongian Linguliform Brachiopods from the Alborz Mountains, Iran
Part of Memoirs of the Association of Australasian Palaeontologists, p. 103-122, 2009
Part of Geological Society, London, Special Publications, p. 23-64, 2009
Part of Geology, p. 171-174, 2008
Part of Earth and Environmental Science Transactions of the Royal Society of Edinburgh, p. 221-232, 2008
The Early Cambrian tommotiid Micrina, a sessile bivalved stem group brachiopod
Part of Biology Letters, p. 724-728, 2008
Middle Cambrian to Lower Ordovician faunas from the Chingiz Mountain Range, central Kazakhstan
Part of Alcheringa, p. 443-463, 2008
- DOI for Middle Cambrian to Lower Ordovician faunas from the Chingiz Mountain Range, central Kazakhstan
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Ordovician ostracod diversity and carbon isotope curve in Gullhögen, Sweden
Part of Acta Palaeontologica Sinica, p. 483-488, 2007
Part of Lethaia, p. 85-96, 2007
Part of GFF, p. 339-348, 2006
Mid Ordovician (Darriwilian) brachiopods of South Kazakhstan
Part of National Museum of Wales Geological Series, p. 145-222, 2006
A spinose stem group brachiopod with pedicle from the Middle Cambrian Burgess Shale
Part of Acta Zoologica, p. 273-290, 2006
Part of Journal of Paleontology, p. 750-759, 2006
Part of Palaeoworld, p. 384-405, 2006
Part of GFF, p. 327-337, 2006
Part of Geobios, p. 125-154, 2006
Part of Memoirs of the Association of Australasian Palaeontologists, p. 1-5, 2006
Early Cambrian brachiopods from North-East Greenland
Part of Palaeontology, p. 325-345, 2005
Part of Journal of Paleontology, p. 884-906, 2005
Part of Geological Magazine, p. 419-439, 2005
Part of GFF, p. 7-16, 2005
The oldest-known metazoan parasite?
Part of Journal of Paleontology, p. 1214-1216, 2004
Part of Geological Magazine, p. 699-715, 2004
Chemico-structure of the organophosphatic shell of siphonotretide brachiopods
Part of Palaeontology, p. 1313-1337, 2004
Early Ordovician brachiopods from the Emanuel Formation, Canning Basin, Western Australia
Part of Memoirs of the Association of Australasian Palaeontologists, p. 113-132, 2004
Early Cambrian lingulate brachiopods from the Shaanxi Province, China
Part of GFF, p. 193-211, 2004
Part of Geological Magazine, p. 31-44, 2003
Understanding linguloid brachiopods: Obolus and Ungula as examples
Part of Camets de Geologie/Notebooks on Geology, 2003
The brachiopod fold: a neglected body plan hypothesis
Part of Palaeontology, p. 59-65, 2003
The Early Cambrian (Botomian) stem group brachiopod Mickwitzia from Northeast Greenland
Part of Acta Palaeontologica Polonica, p. 1-20, 2003
Part of Brigham Young University Geology Studies, p. 23-141, 2003
Lingulate brachiopods from Lower Ordovician (Tremadoc) chacedonite, Holy Cross Mountains, Poland
Part of Acta Palaeontologica Polonica, p. 141-156, 2002
Lingulate brachiopods from the Cambrian-Ordovician boundary beds of Utah
Part of Journal of Paleontology, p. 211-228, 2002
A stem group brachiopod from the Lower Cambrian - support for a Micrina (Halkieriid) ancestry.
Part of Palaeontology, p. 875-882, 2002
Part of Palaentology, p. 845-873, 2002
The Hunneberg Stage (Ordovician) in the area east of S. Petersburg, north-western Russia
Part of Paläontologische Zeitschrift, p. 543-561, 2001
Part of Special Papers in Palaentology, p. 1-180, 2001
Phylogeny and classification: Linguliformea and Craniiformea
Part of The Paleontological Society Papers, p. 11-26, 2001
Small problematic phosphatic sclerites from the Ordovician of Iapetus
Part of Journal of Paleontology, p. 1-8, 2001
Spatial variations in faunal composition, Middle Ordovician, Volkhov Stage, East Baltic
Part of GFF, p. 65-72, 2001
Beachiopoder och konodonter - geologens "svarta lådor"
Part of Geologiskt Forum, p. 3-8, 2000
Redescription of the Ordovician acrotretoid brachiopod Conotreta Walcott, 1889
Part of GFF, p. 313-318, 2000
Part of GFF, p. 367-375, 2000
Cambrian phosphatic brachiopods from the Precordillera of western Argentina
Part of GFF, p. 227-242, 1999
Organophosphatic brachiopods: Patterns of biodiversification and extinction in the early Palaeozoic
Part of Geobios, p. 145-163, 1999
The Late Devonian trematid lingulate brachiopod Schizobolus from Poland
Part of Acta Paleontologica Polonica, p. 335-346, 1999
The diversity and phylogeny of the paterinate brachiopods
Part of Palaeontology, p. 221-262, 1998
Part of Geology, p. 1083-1086, 1998
Late Ordovician and early Silurian Trimerellide brachiopodes from Kazakhstan
Part of Journal of Paleontology, p. 584-598, 1997
Late early Cambrian brachiopods from North Greenland
Part of Palaeontology, p. 337-354, 1997
Part of Journal of Paleontology, p. 577-5883, 1997
Part of Paleontologica Polonica, p. 37-50, 1996
De kalkskaliga brachipodernas tidigaste evolution - nya fynd från Kirgisistan
Part of Geologiskt Forum, p. 10-14, 1996
Distribution of lingulate brachiopods in the early Ordovician of the East Baltic
Part of Danmarks og Grönlands Geologiske Undersögelse, p. 53-56, 1996
Middle Cambrian lingulate brachiopods from Tarbagatay Range, Kazakhstan
Part of Acta Paleontologica Polonica, p. 299-317, 1996
A supra-ordinal classification of the Brachiopoda
Part of Philosophical Transactions of the Royal Society of London. Biological Sciences, p. 1171-1193, 1996
The elkaniide brachiopod Volborthia from the Lower Ordovician of Baltoscandia
Part of Paläontologishes Zeitschrift, p. 213-221, 1995
Phylogenetic analysis and ordinal classification of the Brachiopoda
Part of PALAEONTOLOGY, p. 713-741, 1995
Articles, review/survey
Survival on a soft seafloor: life strategies of brachiopods from the Cambrian Burgess Shale
Part of Earth-Science Reviews, p. 266-287, 2015
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Review of the Ordovician stratigraphy and fauna of the Anarak Region in Central Iran
Part of Acta geologica Polonica, p. 403-435, 2015
- DOI for Review of the Ordovician stratigraphy and fauna of the Anarak Region in Central Iran
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Chapters in book
Kambrosiluren i Västergötland – en geologisk resa genom tid och rum
Part of Sten - en tidsresa, Västergötlands Fornminnesförening, 2017
Biogeography of Ordovician linguliform and craniiform brachiopods
Part of Early Palaeozoic Biogeography and Palaeogeography, p. 117-126, Geological Society of London, 2013
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Part of Brachiopoda, Wiley-Blackwell, 2008
Part of Treatise on Invertebrate Paleontology, p. 2595, The Geological Society of America, University of Kansas, 2007
Part of Treatise on Invertebrate Paleontology, p. 2590-2591, Palaeontological Inst, 2007
Part of Treatise on Invertebrate Paleontology, p. 2592-2594, University of Kansas Press, Lawrence, Kansas, 2007
Part of Treatise on Invertebrate Paleontology, p. 2532-2579, Geological Society of America. and University of Kansas, 2007
Part of The Great Ordovician Biodiversification Event, p. 220-221, Columbia University Press, 2004
Linguliformean and Craniiformean brachiopods
Part of The Great Ordovician Biodiversification Event, Columbia University Press, 2004
Brachipods: Cambrian-Tremadoc precursors to Ordovician radiation events
Part of Palaeobiogeography and Biodiversity Change: the Ordovician and Mesozoic-Cenozoic Radiations, p. 13-23, Geological Society of London, 2002
Part of The Cambrian Biostratigraphy of the Stansbury Basin, South Australia, p. 120-132, Nauka / Interperiodica, 2001
Part of Brachiopods Past and Present, p. 163-176, Department of Palaeontology. The Natural History Museum, 2001
Part of Treatise on Invertebrate Paleontology, Part H (Brachiopoda), p. 30-146, University Press of Kansas, 2000
Part of Treatise on Invertebrate Paleontology, Part H (Brachiopoda) 2 & 3, p. 158-192, Geological Society of America, 2000
Part of Treatise on Invertebrate Paleontology, Part H (Brachiopoda) vol. 2&3, p. 201-208, Geological Society of America, 2000
Part of Treatise on Invertebrate Paleontology, Part H (Brachiopoda) Vol 2 & 3, p. 193-200, Geological Society of America, 2000
Conference papers
Brachiopod Phylogeny in the Cambrian
Part of 8th INTERNATIONAL BRACHIOPOD CONGRESS, p. 56-57, 2018
2012
2012
Early evolution of biomineralization in brachiopods
2012
Brachiopod phylogeny revisited: Using stem groups to flesh out molecular tree topology
2011
Recent palaeobiological and stratigraphical advances from the Cambrian of Estonia
Part of The 2nd Wiman Meeting, p. 5-5, 2011
Part of The Palaeontological Association, 55th Annual Meeting, p. 44-45, 2011
Part of GSA Annual Meeting in Minneapolis, Minnesota USA, p. 543-543, 2011
The Origin of the Brachiopod Body Plan
Part of Internationl Conference on the Cambrian Explosion, p. 541-541, 2009
Part of International Conference on the Cambrian Explosion, Abstract Volume, p. 76-76, 2009
Part of 51st Palaeontological Association Annual Meeting, p. 55, 2007
Part of Rapporter och meddelanden 128, 2007
Part of 51st Palaeontological Association Annual Meeting, p. 37, 2007
Part of The Palaeontological Association, 50th Annual Meeting, 2006, 2006
Part of Lundadagarna i Historisk Geologi och Paleontologi, p. 57, 2005
Part of Lundadagarna i Historisk Geologi och Paleontologi, p. 50, 2005
Shell structure evolution of Cambrian organophosphatic brachiopods
Part of Lundadagarna i Historisk Geologi och Paleontologi, p. 57, 2005
Palaeoecology of a marine impact crater in the Ordovician sea
Part of Palaeontological Association 49th Annual Meeting, 2005
Linguliform and Craniiform Brachiopods from the Ordovician Tvären Crater, Sweden
Part of The 5th International Brachiopod Conference, Copenhagen, Denmark, 2005
A marine impact crater as an Ordovician ecosystem; the Tvären crater
Part of SEPM Research Conference; The sedimentary Record of Meteorite Impacts, 2005
Faunal recovery in the Ordovician Lockne and Tvären craters
Part of Lundadagarna i Historisk Geologi och Paleontologi IX, 2005
Part of Proceedings of the 5th International Brachiopod Congress, Copenhagen, p. 28-29, 2005
Part of Proceedings of the 5th International Brachiopod Congress, Copenhagen, p. 17, 2005
Part of International Subcomission on Cambrian Stratigraphy, 2004
Cambrian brachiopoda of the Rift Valley, Jordan and Israel
2004
Exceptional preservation in the stem group brachiopod Mickwitzia from the Early Cambrian of Nevada
2004
Middle_Upper Ordovician carbon isotopic record from Västergötland (Sweden) and East Baltic
p. 67-68, 2004
p. 987, 2004
"Volkhovian" as a name for the third global stage of the Ordovician System
Part of Ordovician from the Andes, p. 59-40, 2003
The systematic position of the Lower Cambrian brachiopod Heliomedusa Sun and Hou
2003
Depositional sequences in the Ordovician of Baltoscandia
Part of Acta Universitatis Carolinea - Geologica, p. 133-136, 1999
Neodymium isotope composition of Lower Palaeozoic biogenic apatite from Baltoscandia
Part of Acta Universitatis Carolinae - Geologica, p. 511-512, 1999
Early Ordovician (Hunnerberg-Volkhov) facial and faunal changes in the east Baltic
Part of Acta Universitatis Carolinea - Geologica, p. 467-470, 1999
Ordovician patterns of diversification in craniiformean brachiopods
Part of Acta Universitatis Carolinea . Geologica, p. 321-324, 1999
Origins of the Great Ordovician Biodiversification: The record at Lawson Cove, Ibex Area, Utah, USA
Part of Acta Universitatis Carolinea - Geologica, p. 459-462, 1999
Early Paleozoic radiation and classification of organo-phosphatic brachiopods
Part of Proceedings of the third International Brachiopod Congress (Sudbury/Ontario/Canada/2-5 September 1995), p. 117-121, 1996
Part of Ordovician Odyssey: short papers for the Seventh International Symposium on the Ordovician System, p. 319-322, 1995
Part of In J. D. Cooper, M. L. Droser and S.C. Finney (eds.): Ordovician Odyssey: short papers for the Seventh International Symposium on the Ordovician System. Fullerton, California, p. 117-120, 1995